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In spite of Starscream's opinion that Megatron was a "stubborn wreck" for his repeated attempts to sway Orion Pax to his cause, they were successful, and the Decepticons joined forces with Orion's Autobots against Zeta Prime.
After Zeta Prime was defeated, Megatron betrayed Orion and installed himself as ruler of Cybertron, then had the Decepticons hunt down all remaining Autobots.
To cover up his blunder, he drew attention to the spy in their midst, giving Soundwave a cue to reveal himself and stage an escape, distracting Zeta.
His identity evidently unknown to most, Starscream was also positioned as a delegate to the new Senate that was formed by Zeta Prime, though exactly who or where he was masquerading as the delegate of is unclear.
While attending one of Zeta's security sessions, he brought a disguised Soundwave with him to record the Senate's newest plans for how to stop the growing Decepticon menace.
His stubborn refusal to change his ways for the greater good is a constant thorn in the side of his reluctant allies.
Beneath it all, though, Starscream struggles with the ramifications of his morally dubious actions and the growing shadow of his future legacy - a struggle that, in the end, he may not win.